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Publications

peer-reviewed articles

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2024

Schoenemann, B., Hoekstra, H.J.W.M., Horváth, G., Clarkson, E.N.K.
Vision of Trilobites and Polarized Light
In: Horváth, G. (eds) Polarization Vision and Environmental Polarized Light
Springer Series in Vision Research. Springer, Cham (2024)

doi: 10.1007/978-3-031-62863-4_22
https://doi.org/10.1007/978-3-031-62863-4_22

Schoenemann, B., Hughes, N.C., Myrow, P.M.
Eye structure and function in Ameura (Trilobita) from the Upper Carboniferous Minturn Formation at Bond, Colorado, USA
Irish Journal of Earth Sciences 42(1), 299-307 (2024)
doi: https://doi.org/10.1353/ijes.2024.a935035
https://dx.doi.org/10.1353/ijes.2024.a935035

 

2023

Brigitte Schoenemann & Euan N.K. Clarkson
The median eyes of trilobites
Scientific Reports 13, 3917 (2023)
doi: 10.1038/s41598-023-31089-7
https://www.nature.com/articles/s41598-023-31089-7

 

2021

B. Schoenemann, E.N.K. Clarkson, C. Bartels, W. Südkamp, U. Ryck
A 390 million-year-old hyper-compound eye in Devonian phacopid trilobites
Scientific Reports 11, 19505 (2021)
doi: 10.1038/s41598-021-98740-z
https://www.nature.com/articles/s41598-021-98740-z

Brigitte Schoenemann & Euan N.K. Clarkson
Points of view in understanding trilobite eyes
nature communications 12, 2081 (2021)
doi: 10.1038/s41467-021-22227-8
https://www.nature.com/articles/s41467-021-22227-8

Brigitte Schoenemann
An overview on trilobite eyes and their functioning
Arthropod Structure & Development
doi: 10.1016/j.asd.2021.101032
https://www.sciencedirect.com/science/article/pii/S1467803921000062

 

2020

Brigitte Schoenemann & Euan N.K. Clarkson
Insights into a 429-million-year-old compound eye
Scientific Reports
doi: 10.1038/s41598-020-69219-0
https://www.nature.com/articles/s41598-020-69219-0

 

2019

Brigitte Schoenemann, Markus Poschmann & Euan N.K. Clarkson
Insights into the 400 million-year-old eyes of giant sea scorpions (Eurypterida) suggest the structure of Palaeozoic compound eyes.
Scientific Reports
doi: 10.1038/s41598-019-53590-8
https://www.nature.com/articles/s41598-019-53590-8

Oskar Fischer & Brigitte Schoenemann
Why are Bones in Vertebrate Eyes? Morphology, Development and Function of Scleral Ossicles in Vertebrate eyes - a Comparative Study
Journal of Anatomy and Physiological Studies 3(2), 1-26 (2019)

 

2018

Brigitte Schoenemann
Evolution of Eye Reduction and Loss in Trilobites and Some Related Fossil Arthropods
Emerging Science Journal
doi: 10.28991/esj-2018-01151
www.ijournalse.org/index.php/ESJ/article/view/114
www.ijournalse.org/index.php/ESJ/article/download/114/57

 

2017

Brigitte Schoenemann, Helje Pärnaste & Euan N. K. Clarkson
Structure and function of a compound eye, more than half a billion years old
PNAS
doi: 10.1073/pnas.1716824114
www.pnas.org/content/early/2017/11/28/1716824114.full

Brigitte Schoenemann, Euan N. K. Clarkson & Magne Høyberget
Traces of an ancient immune system – how an injured arthropod survived 465 million years ago.
Scientific Reports
7:40330 | DOI:10.1038/srep40330
www.nature.com/articles/srep40330

Sidong Wu, Tao Jiang, Gexiang Zhang, Brigitte Schoenemann, Ferrante Neri, Ming Zhu, Chunguang Bu, Jianda Han & Klaus-Dieter Kuhnert
Artificial compound eye: a survey of the state-of-the-art
Artificial Intelligence Review
DOI: 10.1007/s10462-016-9513-7
link.springer.com/article/10.1007/s10462-016-9513-7

Vizcaíno S. F., Clarkson E.N.K. & Schoenemann B. (Editors)
Form, Function and Palaeobiology.
Special Issue Transactions of the Royal Society of Edinburgh 106 (4) (2017)
https://www.cambridge.org/core/services/aop-cambridge-core/content/view/1BECB5603283FCEC86A6767CA4B705FF/S1755691016000244a.pdf/form_function_and_palaeobiology_preface.pdf

Schoenemann B. & Clarkson E.N.K.
Vision in fossilised eyes.
Special Issue Transactions of the Royal Society of Edinburgh 106 (4), 209-220 (2017)
www.cambridge.org/core/services/aop-cambridge-core/content/view/64E008DCF10488F6C77B07E29BD22293/S1755691016000232a.pdf/vision_in_fossilised_eyes.pdf

 

2016

Vannier J.*, Schoenemann B.* , Gillot T., Charbonnier S. & Clarkson E.N.K.
* contributed equally
Exceptional preservation of eye structure in arthropod visual predators from the Middle Jurassic.
Nature Communications
7:10320 | doi:10.1038/ncomms10320
www.nature.com/ncomms/2016/160119/ncomms10320/full/ncomms10320.html

Schoenemann B (2016)
A Miniscule System of a ‘Sailing Rock’.
J Geol Geophys 5:256 doi: 10.4172/2381-8719.1000256
www.omicsgroup.org/journals/a-miniscule-system-of-a-sailing-rock-2381-8719-1000256.pdf

Poschmann M., Schoenemann B. & McCoy V.
Telltale eyes: The lateral visual systems of Rhenish Lower Devonian eurypterids (Arthropoda, Chelicerata) and their palaeobiological Implications.
Palaeontology, London 59(2), 295–304 (2016)
http://onlinelibrary.wiley.com/enhanced/doi/10.1111/pala.12228

Rust J., Bergmann A., Bartels C., Schoenemann B., Sedlmeier S., & Kühl G.
The Hunsrück biota: A unique window into the ecology of Lower Devonian arthropods.
Arthropod Structure and Development
http://dx.doi.org/10.1016/j.asd.2016.01.004

 

2015

Schoenemann B., Clarkson E.N.K., Horváth G.
Why did the UV-A-induced photoluminescent blue-green glow in trilobite eyes and exoskeletons not cause problems for trilobites?
PeerJ
DOI 107717/peerj.1492
https://peerj.com/articles/1492/

Invited Book Review
Schoenemann B.
Visual Ecology by Cronin, T.W., Johnsen, S., Marshall, N.J., Warrant, E.J., published by Princeton University Press
Visual Ecology. Anat Physiol 5:180. doi: 10.4172/2161-0940.1000180 www.omicsonline.org/open-access/visual-ecology-2161-0940-1000180.pdf

Tanaka G., Schoenemann B., El Hariri K., Ono T., Clarkson E., Maeda H.
Vision in a Middle Ordovician trilobite eye.
Palaeogeography, Palaeoclimatology, Palaeoecology
433, 129–139 (2015)
http://dx.doi.org/10.1016/j.palaeo.2015.05.013

Schoenemann B. & Clarkson E.N.K.
Eyes and vision in the coeval Furongian trilobites Sphaerophthalmus alatus (Boeck, 1938) and Ctenopyge (Mesoctenopyge) tumida Westergård, 1922, from Bornholm, Denmark.
Palaeontology
58(1), 133–140 (2015)
http://onlinelibrary.wiley.com/doi/10.1111/pala.12128/epdf

 

2014

Schoenemann B. & Clarkson E.N.K.
Space, Time and their Relevance to Phylogenetic Analyses (Convergences in Time).
Journal of Geology & Geophysics
DOI: 10.4172/2329-6755.1000178
http://omicsgroup.org/journals/space-time-and-their-relevance-to-phylogenetic-analyses-convergences-in-time-2329-6755-1000178.pdf

Stewart S.E., Clarkson E.N.K., Ahlgren J., Ahlberg P. & Schoenemann B.
Sphenothallus from the Furongian (Cambrian) of Scandinavia.
GFF (Journal of the Geological Society of Sweden)
DOI: 10.1080/11035897.2014.929173
www.tandfonline.com/doi/full/10.1080/11035897.2014.929173

Schoenemann B., Clarkson E.N.K. & Ryck U.
Colour Patterns in Devonian Trilobites.
The Open Palaeontology Journal 8, 113-117 (2014)
http://benthamopen.com/contents/pdf/TOGEOJ/TOGEOJ-8-113.pdf

Schoenemann B., Clarkson E.N.K., Castellani C., Waloszek D., Maas A. & Meyer-Rochow B.V.
Description and Interpretation of the Internal Structure of a Cambrian Crustacean Compound Eye.
Geological Bulletin, Prague 89(2), 311–324 (2014)
www.geology.cz/bulletin/contents/art1429

 

2013

Schoenemann B. & Clarkson E.N.K.
Discovery of some 400 million year-old sensory structures in the compound eyes of trilobites.
Scientific Reports
3:1429 | DOI:10.1038/srep01429
www.nature.com/srep/2013/140313/srep01429/full/srep01429.html

related:
ScienceNOW: Looking a Trilobite in the Eye
(Lizzie Wade)
http://news.sciencemag.org/sciencenow/2013/03/looking-a-trilobite-in-the-eye.html?ref=hp

Schoenemann B.
The eyes of a tiny ‘Orsten’ crustacean – A compound eye at receptor level?
Vision Research Exeter 76, 89-93 (2013)
http://www.sciencedirect.com/science/article/pii/S0042698912003392

Parker A., Schoenemann B., Haug J.T. & Waloszek D.
An unusual cornea from a well-preserved (‘Orsten’) Cambrian compound eye.
Palaeontological Research
, Tokyo 17, 251-260 (2013)
http://www.bioone.org/doi/abs/10.2517/1342-8144-17.3.251

 

2012

Schoenemann B.
The clever strategy of a tiny crustacean eye early in the evolution of vision.
Communicative & Integrative Biology, Austin, Texas 5(2), 175–178 (2012).
DOI: 10.4161/cib.18760
www.ncbi.nlm.nih.gov/pmc/articles/PMC3376057/

Schoenemann B. & Clarkson E.N.K.
At First Sight – Functional Analysis of Lower Cambrian eye systems.
Palaeontographica A, Bonn, Stuttgart 297, 123-149 (2012).
https://www.schweizerbart.de/papers/pala/detail/297/79217/Functional_Analysis_of_Lower_Cambrian_Eye_System

Schoenemann B. & Clarkson E.N.K.
Compound Eyes in the Chengjiang Biota.
Earth and Environmental Sciences of Acta Universitatis Latviensis
, Riga 783, 57-71 (2012).

Schoenemann B. & Clarkson E.N.K.
Insights to Phacopid Eyes.
Earth and Environmental Sciences of Acta Universitatis Latviensis, Riga 783, 72-75 (2012).

Schoenemann B. & Clarkson E.N.K.
The eyes of Leanchoilia.
Lethaia, Copenhagen 45, 524–531 (2012).
http://onlinelibrary.wiley.com/enhanced/doi/10.1111/j.1502-3931.2012.00313.x/

Schoenemann, B., Castellani, C., Clarkson, E. N. K., Haug, J. T., Maas, A., Haug, C. & Waloszek, D.
The sophisticated visual system of a tiny Cambrian crustacean: analysis of a stalked fossil compound eye.
Proceedings of the Royal Society London B 279, 1335–1340 (2012).
DOI: 10.1098/rspb.2011.1888
http://rspb.royalsocietypublishing.org/content/early/2011/10/21/rspb.2011.1888.full.pdf+html

related:
NATIONAL GEOGRAPHIC:Fossil eyes show wraparound three-dimensional vision, half a billion years ago.
(Ed Yong)
http://phenomena.nationalgeographic.com/2011/11/01/fossil-eyes-show-wraparound-three-dimensional-vision-half-a-billion-years-ago/

Castellani, C., Haug, J. T., Haug, C., Maas, A., Schoenemann, B. & Waloszek, D.
Exceptionally well-preserved isolated eyes from Cambrian ‘Orsten’ fossil assemblages of Sweden.
Palaeontology, London 55, 553–566 (2012).
http://onlinelibrary.wiley.com/doi/10.1111/j.1475-4983.2012.01153.x/abstract

 

2011

Schoenemann B. & Clarkson E.N.K.
Eyes and vision in the Chengjiang arthropod Isoxys indicating adaptation to habitat.
Lethaia, Copenhagen 44(2), 223-230 (2011).
http://onlinelibrary.wiley.com/enhanced/doi/10.1111/j.1502-3931.2010.00239.x/

Schoenemann B. & Clarkson E.N.K.
A light guide lens in ancient trilobites?
Transactions of the Royal Society of Edinburgh,  Edinburgh 102, 17-23 (2011).
http://journals.cambridge.org/action/displayAbstract?fromPage=online&aid=8346763&fulltextType=RA&fileId=S1755691011010097

Schoenemann B. & Clarkson E.N.K.
The eyes of Bohemian Trilobites.
Geological Research, Prague 1, 45-50 (2011).

 

2010

Schoenemann B. & Clarkson E.N.K.
The Sophisticated Strategy of a Tiny Eye.
Zitteliana B
, München 29, 92 (2010).
http://epub.ub.uni-muenchen.de/12021/1/zitteliana_2010_b29.pdf
(page 93)

Schoenemann B. & Clarkson E.N.K.
Neue Einblicke in das visuelle System alter Augen – Die Augen von (2010). Phacopiden (Trilobita).
Zitteliana B
, München 29, 92-93

Schoenemann B., Clarkson E.N.K.,  Ahlberg P. & Dies Alvarez M. E.
A Tiny Eye Indicating a Planctonic Trilobite.
Palaeontology, London 53, 695-701 (2010).
http://onlinelibrary.wiley.com/enhanced/doi/10.1111/j.1475-4983.2010.00966.x/

related:
NATIONAL GEOGRAPHIC: Tiny Trilobites Drifted in Cambrian Currents.
(Brian Switek)
http://phenomena.nationalgeographic.com/2010/06/21/tiny-trilobites-drifted-in-cambrian-currents

 

2009

Schoenemann B., Liu J.-n., Shu D.-g., Han J.-a. & Zhang Z.-f.
A Miniscule Optimised Visual System in the Lower Cambrian.
Lethaia, Copenhagen 42, 265–273 (2009).
http://onlinelibrary.wiley.com/enhanced/doi/10.1111/j.1502-3931.2008.00138.x/

 

2008

Schoenemann B.
Analyse früher Augensysteme.
Freiberger Forschungsberichte, Freiberg 15, 85-96 (2008).

Schoenemann B. & Clarkson E.N.K.
Did the trabecula in phacopid lenses act as light-guides?
In: Advances in Trilobite Research (Edts. Rabano I., Gozalo R. & Garcia-Belido D.) 351-355,
Planeta Tierra, Cuadernos Del Museo Geominero No 9, Madrid (2008)

Schoenemann B., Clarkson E.N.K. & Franz A.
Sublensar Capsules in Trilobite Eyes.
In: Advances in Trilobite Research (Edts. Rabano I., Gozalo R. & Garcia-Belido D.) 356-361,
Planeta Tierra, Cuadernos Del Museo Geominero No 9, Madrid (2008)

Schoenemann B., Clarkson E.N.K., Ahlberg P. & Dies Alvarez M. E.
A Furongian Polymerid Planktonic Trilobite.
In: Advances in Trilobite Research (Edts. Rabano I., Gozalo R. & Garcia-Beliodo D.) 361-365,
Planeta Tierra, Cuadernos Del Museo Geominero No 9, Madrid (2008)

 

2007

Schoenemann B.
Trilobite Eyes and a New Type of Neuronal Superpositional Eye in an Ancient System.
Palaeontographica A, Bonn  281, 63-91 (2007).
https://www.schweizerbart.de/papers/pala/detail/281/74286/Trilobite_Eyes_and_a_New_Type_of_Neural_Superposition_Eye_in_an_Ancient_System

 

2006

Schoenemann B.
Cambrian View.
Palaeoworld, Nanjing, Amsterdam 15, 307-314 (2006).
http://www.sciencedirect.com/science/article/pii/S1871174X06000448

 

2005

Schoenemann B.
The Cambrian Sight of View – Suggestions to Investigate Eyes of the Chengjiang Fauna by Comparison with Recent Visual Systems.
Acta Micropalaeontologica Sinica, Nanjing, China 22, 169-170 (2005).

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